ISSN 0042-3211

Supplement to

THE

I VELIGER.

i

I

A Quarterly published by

CALIFORNIA MALACOZOOLOGICAL SOCIETY, INC. Berkeley, California R. Stohler, Founding Editor

Volume 33 January 2, 1990

Supplement 1

^L401

.V4

*

Catalog of Recent and Fossil Molluscan Types in the Santa Barbara Museum of Natural History.

I. Caudofoveata, Polyplacophora, Bivalvia, Scaphopoda, and Cephalopoda

by

PAUL H. SCOTT, F. G. HOCHBERG, and BARRY ROTH

The Veliger (ISSN 0042-3211) is published quarterly on the first day of January, April, July, and October. Membership and subscription information appears on the inside cover. The Veliger is published by tbe California Malacozoological Society, Inc., % Department of Zoology, University of California, Berkeley, CA 94720. Second Class postage paid at Berkeley, CA and additional mailing offices. POSTMASTER: Send address changes to C.M.S., Inc., P.O. Box 9977, Berkeley, CA 94709.

THE VELIGER

Scope of the journal

The Veliger is open to original papers pertaining to any problem concerned with mollusks.

This is meant to make facilities available for publication of original articles from a wide field of endeavor. Papers dealing with anatomical, cytological, distributional, eco- logical, histological, morphological, physiological, taxonomic, evolutionary, etc., aspects of marine, freshwater, or terrestrial mollusks from any region will be considered. Short articles containing descriptions of new species or lesser taxa will be given preferential treatment in the speed of publication provided that arrangements have been made by the author for depositing the holotype with a recognized public Museum. Museum numbers of the type specimen must be included in the manuscript. Type localities must be defined as accurately as possible, with geographical longitudes and latitudes added.

Very short papers, generally not exceeding 500 words, will be published in a column entitled “NOTES, INFORMATION & NEWS”; in this column will also appear notices of meetings, as well as news items that are deemed of interest to our subscribers in general.

Editor-in-Chief

David W. Phillips, 2410 Oakenshield Road, Davis, CA 95616, USA

Editorial Board

Hans Bertsch, National University, Inglewood, California James T. Carlton, University of Oregon

Eugene V. Coan, Research Associate, California Academy of Sciences, San Francisco

J. Wyatt Durham, University of California, Berkeley

William K. Emerson, American Museum of Natural History, New York

Terrence M. Gosliner, California Academy of Sciences, San Francisco

Cadet Hand, University of California, Berkeley

Carole S. Hickman, University of California, Berkeley

David R. Lindberg, University of California, Berkeley

James H. McLean, Los Angeles County Museum of Natural History

Frank A. Pitelka, University of California, Berkeley

Peter U. Rodda, California Academy of Sciences, San Francisco

Clyde F. E. Roper, National Museum of Natural History, Washington

Barry Roth, Santa Barbara Museum of Natural History

Judith Terry Smith, Stanford University

Ralph I. Smith, University of California, Berkeley

Wayne P. Sousa, University of California, Berkeley

T. E. Thompson, University of Bristol, England

Membership and Subscription

Affiliate membership in the California Malacozoological Society is open to persons (no institutional memberships) interested in any aspect of malacology. As an affiliate member, a person may subscribe to The Veliger for US $28.00 (Volume 33), which now includes mailing charges to domestic addresses. There is a one-time membership fee of US $2.00, after payment of which, membership is maintained in good standing by the timely renewal of the subscription; a reinstatement fee of US $3.00 will be required if membership renewals do not reach the Society on or before October 1 preceding the start of the new Volume. If a receipt is required, a self-addressed, stamped envelope (or in the case of foreign members, the envelope and two International Postal Reply coupons) should be included with the membership or subscription request.

The annual subscription rate to The Veliger for libraries and nonmembers is US $56.00 (Volume 33), which now includes mailing charges to domestic addresses.

For subscriptions sent to Canada and Mexico, add US $4.00; for subscriptions sent to addresses outside of North America, add US $8.00, which includes air-expedited delivery.

Memberships and subscriptions are by Volume only (January 1 to October 1) and are payable in advance to California Malacozoological Society, Inc. Single copies of an issue are US $25.00 plus postage.

Send all business correspondence, including subscription orders, membership applications, payments for them, changes of address, to: C.M.S., Inc., Post Office Box 9911, Berkeley, CA 94709, USA.

Send manuscripts, proofs, books for review, and correspondence regarding editorial matters to: Dr. David W. Phillips, Editor, 2410 Oakenshield Road, Davis, CA 95616, USA.

THIS PUBLICATION IS PRINTED ON ACID-FREE PAPER.

The Veliger 33(Suppl. l):l-27 (January 2, 1990)

THE VELIGER

© CMS, Inc., 1990

Catalog of Recent and Fossil Molluscan Types in the Santa Barbara Museum of Natural History.

I. Caudofoveata, Polyplacophora, Bivalvia, Scaphopoda, and Cephalopoda

by

PAUL H. SCOTT, F. G. HOCHBERG, and BARRY ROTH

Department of Invertebrate Zoology, Santa Barbara Museum of Natural History,

2559 Puesta del Sol, Santa Barbara, California 93105, USA

Abstract. The non-gastropod molluscan types currently housed in the Department of Invertebrate Zoology at the Santa Barbara Museum are listed. Three hundred seventeen type lots are reported, representing 211 recent species and 9 species originally described as fossils. Each type lot recorded includes a complete citation, type locality, and the current type status of the specimens. An author index and alphabetic index are provided. Lectotype designations are made for three species: Mopalia chacei Berry, 1919; Placiphorella pacifica Berry, 1919; and Venus fordii Yates, 1890.

INTRODUCTION

In the past five years the Santa Barbara Museum of Nat- ural History has added approximately 1400 primary mol- luscan type lots to its holdings. The majority of these specimens were contained in the extensive mollusk collec- tion of the late S. Stillman Berry, which the museum received in 1985 (Hochberg, 1985). Many of the types represent species described by Berry (for a complete listing of Berry’s taxa see Sweeney & Roper, 1984). Also in- cluded are a number of important species from the eastern Pacific Ocean that Berry obtained through exchange with the United States National Museum of Natural History, the Harvard University Museum of Comparative Zoology, and the Philadelphia Academy of Natural Sciences. Types acquired in exchange by Berry include species described by P. P. Carpenter, C. B. Adams, VV. H. Dali, and many other authors.

The Department of Invertebrate Zoology at the Santa Barbara Museum of Natural History was formally estab- lished in 1958, although the museum was founded in 1916. A number of important mollusk collections, donated in the early 1940s, form the nucleus around which the depart- ment was organized. The current shelled mollusk collec- tion contains about 180,000 species lots. With the addition of cephalopods from the Allan Hancock Foundation our

current holdings of octopuses and squids totals over 7000 species lots.

A number of mollusk paratypes and hypotypes were destroyed in a fire at the museum in 1962. SPHON (1962) detailed the types lost in the fire and later (SPHON, 1966) published an updated list of types housed in the depart- ment.

Several works on the molluscan taxa of selected authors were consulted during the compilation of this catalog. Hertz (1984) illustrated the types of S. S. Berry that were described in his Leaflets in Malacology. The species de- scribed by H. N. Lowe, and descriptions Jointly authored by H. A. Pilsbry & Lowe, are listed in Hertz (1986), along with illustrations of previously unfigured types. Names introduced by H. A. Pilsbry are listed in Clench & Turner (1962). Turner (1956) listed and illustrated the eastern Pacific species of C. B. Adams, and included many lectotype designations. W. H. Dali’s taxa are listed in Boss et al. (1968). Palmer (1958, 1963) and Keen (1968) discussed the eastern Pacific taxa of P. P. Carpenter and provide numerous type illustrations.

FORMAT

This catalog lists primary types (holotypes, paratypes, syn- types, neotypes, lectotypes, and paralectotypes) of recent

Page 2

The Veliger, Vol. 33, Suppl. 1

and fossil mollusks in the classes Caudofoveata, Polypla- cophora, Bivalvia, Scaphopoda, and Cephalopoda. Records of the gastropod types will be included in a future catalog. Types not regulated by the International Code of Zoological Xomenclature (Ride et al., 1985) (hereafter ICZX), in- cluding hypotypes and plaster casts of types, are excluded. The catalog is arranged systematically by class, subclass or order, and family. Species-group names are arranged alphabetically within each family. Two indices are pro- vided: an author index that lists all taxa described by a given author, and an alphabetic index by species-group name.

The format for each species-group name is as follows:

species. Genus {Subgenus)

Author, date: Original citation.

Type category: Catalog number (number of specimens, preservative); last catalog number if any.

Locality as published in original description.

Remarks: Lectotype information, specimen condition, other previous catalog numbers, etc.

The following general guidelines are used:

Genus-group names are as originally proposed and do not include new combinations.

Family-group names are those currently used in the literature. Bernard (1983) w'as the primary source for families of the Bivalvia, Kaas & V'an Belle (1980) and Van Belle (1981) for the Polyplacophora, Voss (1977) and Cl.arke (1988) for the Cephalopoda, and Boss (1982) for the Caudofoveata and Scaphopoda.

The original citation includes only figures and plates that illustrate actual specimens in the collection of the Santa Barbara Museum of Natural History.

The type series is the most difficult category to delin- eate. The ICZX (Ride et ai, 1985) was used as our guide- line. We have tried to use the code conservatively to avoid subsequent confusion with the status of a type series. Each type lot was categorized using the following definitions.

Holotype the single specimen designated by the au- thor to be the name-bearing type in the original pub- lication. In a few instances we have demonstrated that a species-group taxon was based on a single specimen and have designated the holotype by monotypy [7CZA’ .\rt. 73a (ii)].

Paratypes all specimens remaining in a type series after a holotype has been designated, or other speci- mens designated as paratypes in the original descrip- tion.

Syntypes all specimens in a type series in w'hich nei- ther a holotype nor a lectotype has been designated.

Lectotype a specimen in a type series that has been designated by a subsequent author as the name-bear- ing type. This includes specimens that have been in- advertently referred to in print as the “holotype” or “the type” {ICZX Art. 74b). We have attempted to trace the current status of each species group and

include lectotype designations, although some desig- nations may have been overlooked.

Lectotype (herein) In several instances S. S. Berry neglected to designate types in the original publication, but labeled his specimen as the “Type” or “Holotype” and entered them as such in his collection catalog. We feel the authors intent was clear for these type spec- imens and have chosen to designate lectotypes in ac- cordance with ICZX Art. 74a. We have also desig- nated a lectotype for one species described by L. G. Yates.

Paralectotypes all specimens remaining in the type series after a lectotype has been designated.

The number of specimens in each type lot is stated and, where applicable, information on incomplete speci- mens (single valves or plates, etc.) is included. Bivalves are listed as a matched pair of valves (pair) or as single un- matched valves (valves). In a few cases a dash (-) in this section indicates that the specimen is represented by a small fragment. Such cases are further mentioned in the remarks section. The current preservative is either a dried shell body (dry), 70% ethyl alcohol (70% EtOH), or balsam.

The immediately previous catalog number is given when available. Other previous numbers are given in the remarks section. S. S. Berry frequently gave his chiton specimens two numbers. The first number is his general catalog number, the second [in brackets] is a chiton catalog number.

The locality is as published in the original description and may include additional information from the specimen label. We have included current higher geographic cate- gories (country, state, province, etc.) to maintain a consis- tent format. In addition, if station numbers were given in the original description {e.g., United States Fish Commis- sion Stations) we have included the latitude and longitude, or bearing of the station when available. Localities in the United States begin with the state in uppercase; localities in other countries or oceans begin with the country or ocean in uppercase. Anglicized names have been changed to their proper place name {e.g., Angeles Bay is changed to Bahia de los Angeles). If a place name is no longer in common use we have provided the currently accepted name in brackets. Metric equivalents have been included in brackets if non-metric units of measurement were included in the original description or label.

The remarks include reference to lectotype designations and anyAmportant data not mentioned in the above cat- egories. When available we have included the current re- pository of the holotypes and lectotypes for those taxa that the Santa Barbara Museum of Natural History houses only paratypes or paralectotypes.

ABBREVIATIONS

Standard abbreviations as recommended in the Council of Biology Editors Style Manual (Huth et al., 1978) have been

P. H. Scott et ai, 1990

Page 3

used throughout the text. In addition the following col- lection abbreviations have been used.

AHF Allan Hancock Foundation Cephalopod Type Collection

AXSP Academy of Natural Sciences of Philadelphia CAS California Academy of Sciences, San Francisco CMNH Carnegie Museum of Natural History, Pitts- burgh

LACM Los Angeles County Museum of Natural His- tory

MCZ Museum of Comparative Zoology, Harvard University

SBMNH Santa Barbara Museum of Natural History SDNHM San Diego Natural History Museum SSB S. Stillman Berry Collection USFC United States Fish Commission USNM United States National Museum of Natural History

ACKNOWLEDGMENTS

Gene Coan and Henry Chaney provided many helpful comments on sections of this manuscript. Bill Emerson (American Museum of Natural History, New York) as- sisted with the determination of type status for the scaph- opods. George Kennedy (Los Angeles County Museum of Natural History) provided current data on the fossil lo- calities. Henry Chaney photographed the chiton lectotypes. The following individuals and institutions provided infor- mation on their type holdings: Robert Van Syoc and Mi- chael Kellogg (California Academy of Sciences); Diane Tyler and Mike Sweeney (U.S. National Museum of Nat- ural History); Clif Coney and Gale Sphon (Los Angeles County Museum of Natural History).

LITERATURE CITED

Bernard, F. R. 1983. Catalogue of the living Bivalvia of the eastern Pacific Ocean: Bering Strait to Cape Horn. Canadian Special Publications of Fisheries and Aquatic Sciences 61: viii + 102 pp.

Boss, K. J. 1982. Mollusca. Pp. 945-1166. In: S. P. Parker (ed.). Synopsis and classification of living organisms. 1. McGraw-Hill Book Co.; New York, New York, viii +1166 pp.; 87 pis.

Boss, K. J., J. Rosewater & F. Ruhoff. 1968. The zoo- logical taxa of William Healey Dali. United States National Museum Bulletin 287:vi + 427 pp.

Clarke, M. R. 1988. Evolution of recent cephalopods a brief review. Pp. 331-340. In: M. R. Clarke & E. R. Trueman (eds.). The Mollusca. \’ol. 12. Paleontology and neontology of cephalopods. .\cademic Press: .San Diego. California. Clench. \V. J. 1962. New names introduced by H. .\. Pilsbry in the Mollusca and Crustacea, .\cademy of Natural Sciences of Philadelphia, Special Publication 4:vi -^218 pp.

Hertz. C. M. 1984. Illustrations of the types named by S. Stillman Berry in his “Leaflets in Malacology.” Festivus 1 5(.Suppl.);l-42; 92 figs.

Hertz. C.M. 1986. Herbert Nelson Lowe 1880-1936 a gifted

amateur conchologist from southern California. Festivus 18(3):26-43; 17 figs.

Hochberg, F. G. 1985. Museum acquires S. Stillman Berry mollusk collection and library. Santa Barbara Museum of Natural History Bulletin 83(April):l.

Huth, E. j. et al. 1978. Council of Biology Editors style man- ual: a guide for authors, editors, and publishers in the bio- logical sciences. 4th ed. Council of Biology Editors. 265 pp.

Kaas, P. & R. Van Belle. 1980. Catalogue of living chitons. Dr. W. Backhuys: Rotterdam, The Netherlands. 144 pp.

Keen, A. M. 1968. West American mollusk types at the British Museum (Natural History). IV. Carpenter’s Mazatlan Collection. Veliger 10(4):389-439; pis. 55-59.

Palmer, K. V. W. 1958. Type specimens of marine Mollusca described by P. P. Carpenter from the West Coast (.San Diego to British Columbia). Geological Society of America, Memoir 76:viii + 376 pp.; 35 pis.

Palmer, K. V. W. 1963. Type specimens of marine Mollusca described by P. P. Carpenter from the west coast of Mexico and Panama. Bulletins of American Paleontology 46(211): 285-408; pis. 58-70.

Ride. W. D. L. el al. (eds.). 1985. International code of zoo-

logical nomenclature adopted by the XX General .Assembly of the International Union of Biological Sciences. 3rd ed. University of California Press: Berkeley, California. 338 pp.

Sphon, G. G. 1962. Destroyed type material. Veliger 5(1 ):57.

Sphon, G. G. 1966. Material contained in the molluscan type collection of the Santa Barbara Museum of Natural History. Veliger 9(2):244-246.

Sweeney, M. J. & C. F. E. Roper. 1984. The zoological taxa and bibliography of S. Stillman Berry. .American Malacolog- ical Bulletin 3(l):63-82.

Turner, R. D. 1956. The eastern Pacific marine mollusks described by C. B. .Adams. Harvard University, Museum of Comparative Zoology, Occasional Papers on Mollusks 2(20): 21-136; pis. 5-21 .

Van Belle, R. .A. 1981. Catalogue of fossil chitons. Dr. W. Backhuys: Uitgever, Rotterdam, The Netherlands. 84 pp.

Voss, G. L. 1977. Classification of recent cephalopods. Pp. 575-579. In: M. Nixon & J. B. Messenger (eds.). The biology of cephalopods. .Academic Press: New York, New York.

TYPE CATALOG CLASS CAUDOFOVEATA

Chaetodermatidae

montereyensis, Chaetoderma

Heath, 1911: Mem. Mus. Comp. Zool. 45(l):61-62. .Syntype: .SBMNH 35034 (1, dry).

California, Monterey Co., Monterey Bay, off Pt. Pinos Light [bearing S 10°W, 9.9 mi (15.9 km)]; 228-213 fm [417-390 mj; USFC Sta. 4524.

Remarks: Specimen dried, previously preserved in fluid (pos- sibly alcohol).

CLASS POLYPLACOPHORA ORDER NEOLORICATA

Leptochitontdae

abbreviatus, Lepidopleurus halistreptus

Dall, 1908: Bull. Mus. Comp. Zool. 43(6):354.

Syntypes: SBMNH 34461 (3, dry); previous number S.SB 1 4556.

Page 4

The Veliger, Vol. 33, Suppl. 1

Mexico, Guerrero, off Acapulco; 16°32'N, 99°48'\V; 493 fm [902 m]; green mud; USFC Sta. 3417.

Remarks: .\11 specimens articulated, ex USNM 223498.

arnbustus, Lepidopleurus

D.\LL. 1919; Proc. U.S. Natl. Mus. 55(2283):499-500.

Syntype: .SBMNH 34462 (1, dry); previous number SSB 14557.

Caliform.a, Santa Barbara Co., Santa Barbara Channel; 20- 40 fm [37-73 m].

Remarks: Specimen articulated, ex USNM 274120.

clarki, Leptochiton

Berry, 1922: Proc. Calif. .\cad. -Sci. (4)1 1(18):412, 427-430, pi. 1, fig. 10, text figs. 1-4.

Holotype: SBMNH 34397 (1, dry); previous number SSB 3987 [605].

Californi.a, Los Angeles Co., Santa Monica [=Los .Angeles], Long Wharf Canyon [Potrero Canyon]; San Pedro Pleis- tocene [Palos \’erdes Sand or equivalent, upper Pleistocene].

Remarks: 1 intermediate valve in 2 pieces. .S. .S. Berry originally described the type locality as Santa Monica; however, the locality is now within the city limits of Los .\ngeles.

jarallonis, Lepidopleurus

Dale, 1902: Proc. U.S. Natl. Mus. 24(1264):557.

Syntype: SBMNH 34458 (-, dry); previous number SSB 27978.

Californi.a, San Francisco Co., off Farallon Ids.; 37°23'N, 123°08'W; 391 fm [715 m]; clay; USFC Sta. 3104.

Remarks: S. S. Berry label “fragment of right side of valve 7 of holotype.” ex USNM 109025.

heathi. Lepidopleurus (Xiphiozona)

Berry. 1919: Lorquinia 2(6):45.

Lectotype; SBMNH 34380 (1, dry); previous number SSB 3513 [123].

Paralectotype: SBMNH 34381 (1, dry); previous number SSB [124a].

Paralectotypes: SBMNH 34382 (5, 70% EtOH); previous number SSB [ 1 24].

California. Monterey Co., off Monterey; 15 fm [27 m[.

Remarks: Lectotype designated: Berry. S. S. 1919. Proc. Cal- ifornia .Acad. Sci. (4)9(1 );6-8; lectotype illustrated in pi. 1, figs. 1, 2; SBMNH 34381 illustrated in pi. 2, figs. 1-6.

opaciis, Lepidopleurus

Dale, 1908: Bull. Mus. Comp. Zool. 43(6):218. 354-355.

Syntype: SBMNH 34460 ( 1 , dry); previous number .SSB 23883.

Southeast Pacific Ocean, between the Galapagos Ids. and mainland Peru; 04°33'S, 87°42'30"W; 2005 fm [3667 m[; ooze; USFC .Sta. 4647.

Remarks: 1 valve, ex USNM 110664.

H.ANLEYIDAE

spicata. Hanleya

Berry. 1919: Proc. Calif. .Acad. .Sci. (4)9(1):8-10. pi. 1, fig. 3.

Holotype: .SBMNH 34396 (1, balsam); previous number SSB 4102 [63].

California. Monterey Co., Monterey Bay. ledge off Pt. Pinos; on a fragment of rock; “depth probably 200 ft” [61 m].

Remarks: .Specimen cleared and mounted on a slide in balsam; cover slip is cracked, obscuring \ iew of specimen.

ISCHNOCHITONIDAE

aelhotiiis. Ischnochilon (Slenoplax)

Dale. 1919: Proc. U.S. Natl. .Mus. 55(2283):507-508.

Syntype: SB.MNH 34464 (-, dry); previous number .S.SB 27979.

Mexico. Baja California Sur. near Cabo .San Lucas;

24‘’11’45"N, 109“55'W; 10 fm [18 m]; shell bottom; USFC .Sta. 2827.

Remarks: S. S. Berry label “shred of holotype.” ex USNM 218814.

amabilis, Ischnochiton (Lepidozona)

Berry, 1917; Proc. U.S. Natl. Mus. 54(2223):6-8, pi. 4, fig. 5, pi. 5, figs. 1-4.

Paratvpes: SBMNH 34377 (2, dry); previous number SSB [113].

Japan, Tsugaru-Heiya Pref., off Cape Tsiuka; 40°33'50"N, 140°36'45"E; 47 fm [86 m]; bottom of sand, shells, and coarse gravel; USFC Sta. 4808.

Remarks: One specimen articulated on girdle, 1 specimen dis- articulated, without girdle. Holotype USNM 215630.

asthenes, Ischnochiton {Lepidozona)

Berry, 1919: Lorquinia 2(6):47.

Lectotype; SBMNH 34389 (1, dry); previous number SSB

3913 [466].

Paralectotype; SBMNH 34390 (1, dry); previous number SSB

3914 [471a].

Paralectotypes: SBMNH 34391 (15, dry); previous number .SSB 3914 [471].

Paralectotypes: SBMNH 34392 (5, dry); previous number SSB [472].

California, Los .Angeles Co., White’s Point; under stones at low tide.

Remarks: Lectotype designated: Berry, S. S. 1919. Proc. Calif. •Acad. Sci. (4)9(1):18-21; SBMNH 34390 illustrated in pi. 8, figs. 1, 2. Lectotype specimen articulated,

californiensis, Ischnochiton {Lepidozona)

Berry, 1931: Proc. Malacol. Soc. London 19(5):255-258, pi. 29, fig. 1 .

Holotype: SBMNH 34407 (1, dry); previous number SSB 5226 [1699].

Paratypes: SBMNH 34408 (10, dry); previous number SSB 3119 [65].

California, San Diego Co., La Jolla, rocky shore near Scripps Institution of Oceanography; between tides.

Remarks: .All specimens articulated. Holotype is illustrated in pi. 29, fig. 1 .

circumsenta, Stenoplax

Berry, 1956: Leafl. Malacol. 1(13):72.

Holotype: SBMNH 34425 (1, dry); previous number SSB 13602 [2227].

Paratype: SBMNH 34491 (1, dry); previous number SSB 13627 [2228].

Mexico, Baja California Sur, Scammon’s Lagoon [Laguna Ojo de Liebre], sand flats W of Isla Concha; under shells of Pecten circularis.

Remarks: Holotype articulated, paratype partially disarticu- lated.

connellyi. Callistochiton

Willett, 1937: Nautilus 51:25-26.

Paratypes: SB.MNH 34478 (4, dry); previous number SSB 10411.

Mexico. Baja Cialifornia Norte, near Ensenada. .Arbolitos Point jsouth of Punta Banda?].

Remarks: .Ml specimens articulated, ex Cionnelly collection. Holotype L.ACiM 1048. crossota. Xuttallina

Berry, 1956: Leafl. Malacol. l(13):71-72.

Holotype: SBMNH 34423 (1, dry); previous number SSB 15133.

P. H. Scott et al., 1990

Page 5

Paratypes: SBMNH 34424 (13, dry); previous number SSB 14966.

Mexico, Sonora, Puerto Penasco, \V end of long bight; under stones in tide pools on reef.

Remarks: Holotype articulated; paratypes 12 articulated specimens, 1 disarticulated specimen.

euryplax, Chaetopleura (Pallochiton)

Berry, 1945: .\mer. Midi. Natur. 34(2):491-492, text figs. 1-9.

Holotype: SBMNH 34409 (1, dry); previous number SSB 8943 [2095].

Paratypes: SBMNH 34410 (2, dry); previous number SSB 8564.

Mexico, Sonora, Bahia de Adair; taken alive under stones at low tide.

Remarks: Holotype 8 disarticulated valves and dried girdle; paratypes 1 disarticulated specimen with radula, 1 artic- ulated specimen with radula, 1 unmatched radula.

fackenthallae, Cyanoplax

Berry, 1919: Lorquinia 2(6):45.

Holotype (monotypy): SBMNH 34383 ( 1 , dry); previous num- ber SSB 3907 [441].

California, Monterey Co., Pacific Grove.

Remarks: 8 disarticulated valves (No. 7 broken) and dried girdle. S. S. Berry did not designate a holotype in the original description but his card catalog indicates only one specimen in the type lot. No other specimens that were collected before tbe date of the species description were located in his col- lection.

ferminicus, Callistochiton decoratus

Berry, 1922: Proc. Calif. Acad. Sci. (4)1 1(18):409, 483, pi. 14, figs. 7, 8.

Holotype: SBMNH 34401 (1, dry); previous number SSB 4571 [1040].

Paratype: SBMNH 34402 ( 1 , dry); previous number SSB 4572 [1095].

California, Los Angeles Co., Pt. Fermin, “Chiton Bed”; low- er San Pedro Pleistocene [upper (?) Pleistocene].

Remarks: Each lot has 1 intermediate valve. The type locality is the second terrace of \V OODRING et al. ( 1 946, U.S. Geologic Survey Professional Paper 207).

fisheri, Callistochiton

Dall, 1919: Proc. U.S. Natl. Mus. 55(2283):512.

Syntype: SBMNH 34467 (1, dry); previous number SSB 14561.

Alaska, Aleutian Ids., Tanaga Id., Glory of Russia Bay.

Remarks: Specimen partially disarticulated, ex USNM 1 10353.

gallina, Ischnochiton {Lepidozona)

Berry, 1925: Proc. Malacol. Soc. London 16(5):228-229, pi. 1 1 , figs. 1 , 2.

Holotype: SBMNH 34404 (1, dry); previous number SSB 4898 [757].

California, Los Angeles Co., ofT White’s Point.

Remarks: Specimen articulated, valve 2 is cracked.

golischi, Ischnochiton {Lepidozona)

Berry, 1919: Lorquinia 2(6):47.

Lectotype: SBMNH 34395 (1, dry); previous number SSB 4093 [1068].

California, Los Angeles Co., off Santa Monica; 100 fm [183 m]; from coral.

Remarks: Lectotype designated: Berry, S. S. 1925. Proc. Mal- acol. Soc. London 21(5):230, pi. 11, figs. 3, 4.

heathiana, Stenoplax {Stenoradsia)

Berry, 1946: Proc. Malacol. Soc. London 26(6): 161 -164, pi. 4, figs. 7, 9, text figs. 1-6.

Paratypes: SBMNH 34415 (4, dry); previous number .SSB 885 [294].

California. Monterey Co., Pacific Grove, shore of Monterey Bay; between tides.

Paratypes: SBMNH 34416 (2, dry); previous number SSB 4728 [1218].

Paratype: SBMNH 34417 (1, dry & 70% EtOH); previous number SSB 8572 [131].

Paratype: SBMNH 34418 (1, dry & 70% EtOH); previous number SSB 8572c [131c].

Paratype: SBMNH 34419 (1, 70% EtOH); previous number SSB [2097].

California, Monterey Co., Pacific Grove, on the shore be- tween the “Third Beach” and the Marine Laboratory; be- tween tides.

Remarks: .SBMNH 34416 illustrated in pi. 4, figs. 7, 9. .SBMNH 34417 illustrated in text figs. 1-6. SBMNH 34418 articulated 7 valve specimen, body in alcohol. .SBMNH 34419 body only without valves or girdle. Ho- lotype C.-\S 064459.

histrio, Stenoplax

Berry, 1945: Amer. Midi. Natur. 34(2):493-495.

Holotype: SBMNH 34411 (1 valve, dry); previous number SSB 1 1982 [1902].

Paratypes: SBMNH 34412 (4, dry & 70% EtOH); previous number SSB 7116 [1901].

Mexico, Baja California Sur, Mulege, El Gallo [Puma Gal- lito?].

Remarks: Holotype only valve 4, remaining valves of holotype at California .Ucademy of Sciences (CAS 029671); para- types— 3 articulated specimens, 1 disarticulated specimen, 3 bodies in alcohol.

imitator. Eudoxochiton

Iredale, 1914: Proc. Malacol. Soc. London 11(1):30-31.

Syntype: SBMNH 34475 ( 1 , dry); previous number SSB 5518.

New Zealand, Kermadec Ids., Sunday Id.

Remarks: .Specimen articulated.

inefficax, Lepidozona

Berry, 1963: Leafl. Malacol. 1(22): 138.

Paratype: SBMNH 34447 (1, dry); previous number SSB 28711.

Mexico, Baja California Norte, Bahia Sebastian Vizcaino; 28°26.5'N, 114°36'VV; 55-57 fm [101-104 m]; on the bra- chiopod Terebratalia obsoleta; trawler “Alaska.”

Remarks: This specimen is mentioned in the original descrip- tion. and was collected with the holotype. The holotype, apparently lost (Hertz, 1 984), is not at the California Acad- emy of Sciences (M. Kellogg, 1988, pers. comm.).

interfossa, Ischnochiton {Lepidozona)

Berry, 1917: Proc. U.S. Natl. Mus. 54(2223):8-10, pi. 3, figs. 8, 9, pi. 5, figs. 5-8.

Paratypes: SBMNH 34378 (2, dry); previous number SSB [114].

Japan, Tsugaru-Heiya Pref., off Cape Tsiuka; 41°35'50"N, 140°36'45"E; 47 fm [86 m]; bottom of sand, shells, and coarse gravel; USFC Sta. 4808.

Remarks: Holotype USNM 215631.

intermedins, Ischnochiton

Hedley & Hull, 1912: Proc. Linnean Soc. New S. Wales 37(2):274-275.

Page 6

The Veliger, Vol. 33, Suppl. 1

Syntypes: SBMNH 34471 (3, dry); previous number SSB 5237.

Norfolk Island.

Remarks: .All specimens articulated.

isoglypla, Stenoplax

Berry, 1956: Leafl. Malacol. l(13):72-73.

Holotype: SBMNH 34426 (1, dry); previous number SSB 19160 [2226].

Peru. Isabel Id.

Remarks: 6 disarticulated valves, 1 dried body. Portions of the holotype distributed by S. S. Berry; valve No. 6 to USNM 612205, and 1 valve to CAS 043940. The authors have been unable to locate the type locality of Isabel Island.

keepiana, Lepidochitona

Berry, 1948: Leafl. Malacol. 1(4):14-15.

Holotype: SBMNH 34420 (1, dry); previous number SSB 11992 [2146].

Paratypes: SBMNH 34421 (7, dry); previous number SSB 3978 [868].

CaLIFOR.nia, Orange Co., Newport Bay.

Remarks: Holotype articulated; paratypes 4 articulated spec- imens, 2 partially disarticulated specimens, 1 disarticulated specimen.

laurae. Dendrochiton

Berry, 1963: Leafl. Malacol. 1(22):136.

Holotype: SBMNH 34439 (1, dry); previous number SSB 29878.

Mexico, Baja California Norte, Bahia de los .Angeles.

Remarks: Specimen articulated.

lincolnensis, Ischnochiton atkmsom

■\SHBY, 1920: Trans. Roy. Soc. So. Australia 44:275-277.

Syntype: SBMNH 34455 (1, dry); previous number SSB 6723.

Australia, South Australia, Port Lincoln.

Remarks: Specimen articulated.

lioplax, Oligochiton

Berry, 1922: Proc. Calif. Acad. Sci. (4)1 1(18):405, 431-433, pi. 1, figs. 1-6.

Paratype: SBMNH 34398 ( 1 , dry); previous number SSB 5052 [606].

Canada, British Columbia, Vancouver Id., VV of Otter Point, between mouths of Muir and Kirby creeks; from conglom- erates and sandstones along the sea cliff; Oligocene [lower Miocene, Juanian Stage], Sooke Formation; Stanford Uni- versity (North Pacific) locality N.P. 129.

Remarks: 1 head valve attached to matrix. Holotype CAS 61805.46.

lirulatus, Dendrochiton

Berry, 1963: Leafl. Malacol. 1(22): 136- 137.

Holotype: SBMNH 34440 (1, dry); previous number SSB 29879.

Paratypes: .SBMNH 34442 (7, dry); previous number SSB 29826.

Paratypes: SBMNH 34443 (12, dry); previous number SSB 29833.

Mexico, Baja California Norte, N of San Felipe, Ensenada Blanca; under stones in low intertidal rubble.

Paratypes: SBMNH 34441 (2, dry); previous number SSB 25452.

Mexico, Baja California Norte, N cove of San Felipe.

Remarks: .311 specimens articulated except SBMNH 34441, which is partially disarticulated.

lobium, Basiliochiton

Berry, 1925: Proc. Acad. Natur. Sci. Philadelphia 77:27-28, pi. 2, figs. 1, 2.

Holotype: SBMNH 34403 (1, dry); previous number SSB 4908 [1353].

California, San Diego Co., La Jolla, tide pool in the “Devil’s Slide.”

Remarks: Specimen articulated but all valves cracked.

magdalena, Xuttalina

Dall, 1919: Proc. U.S. Natl. Mus. 55(2283):502.

Syntype: .SBMNH 34463 (1, dry); previous number SSB 14558.

Mexico. Baja California Sur, Bahia Magdalena.

Remarks: Specimen articulated, ex USNM 217924.

mixta. Tonicia

Dall, 1919: Proc. U.S. Natl. Mus. 55(2283):515-516.

Syntype: SBMNH 34468 (1, dry); previous number SSB 14562.

Mexico, Baja California Norte, Bahia de los Angeles.

Remarks: Specimen articulated.

nipponica, Ischnochiton: Berry, 1918 (see Ischnochiton pilsbry- anus)

pella. Lepidozona

Berry, 1963: Leafl. Malacol. 1(22):137-138.

Holotype: SBMNH 34444 (1, dry); previous number SSB 25471.

Paratype: SBMNH 34445 (1, dry); previous number SSB 29827.

Mexico, Baja California Norte, San Felipe; under the cleaner stones on a very muddy rocky shore, low mid-tide.

Paratypes: SBMNH 34446 (14, dry); previous number SSB 29827.

Mexico, Baja California Norte, San Felipe, below El Faro, under muddy rocks, low mid-tide.

Remarks: .<311 specimens articulated.

perplexus, Eudoxochiton

Iredale, 1914: Proc. Malacol. Soc. London 11(1):29.

Syntype: SBMNH 34474 (1, dry); previous number SSB 5230.

New Zealand, Kermadec Ids., Sunday Id.

Remarks: Specimen articulated.

pilsbryanus, Ischnochiton {Lepidozona)

Berry, 1917: Proc. U.S. Natl. Mus. 54(2223):10-12, pi. 7, figs. 1-7.

Paratype: SBMNH 34379 (1, dry); previous number SSB [119].

Japan, Oshima-Hauto Pref., Hokkaido, off Cape Shirakami; 4I°17'20"N, 140°07'\V; 195 fm [357 m]; bottom of fine gray sand, USFC Sta. 4810.

Remarks: Junior homonym of Ischnochiton (Stenochiton) pils- bryanus Bendall, 1897; renamed Ischnochiton {Lepidozona) nipponica Berry, 1918; Nautilus 3 1(4): 144. Holotype USNM 215632.

psaltes, Dendrochiton

Berry, 1963: Leafl. Malacol. 1(22):135.

Holotype: SBMNH 34435 (1, dry); previous number SSB 29877.

Paratypes: SBMNH 34436 (26, dry).

Paratype: SBMNH 34437 (1, dry).

Paratype: SBMNH 34438 (1, dry).

California, San Diego Co., Mission Bay, near entrance.

Remarks: All specimens articulated. SBMNH 34438 is a 7-valved specimen.

saccharina, Tonicella

Dall, 1878: Proc. U.S. Natl. Mus. 1(1):2.

Paralectotype: SBMNH 34457 (1, dry); previous number SSB 14558.

■Alaska, Shumagin Ids., Yukon Harbor; 13 fm [24 m[.

P. H. Scott et al., 1990

Page 7

Remarks: Specimen articulated. Lectotype designated: Ferreira, A. J. 1982. Veliger 25(2):1 14-1 15; USNM 30914.

sanctaemonicae , Ischnochiton (Lepidozona)

Berry, 1922: Proc. Calif. Acad. Sci. (4)1 1(18):412, 478-480, pi. 12, figs. 1-9.

Holotype: SBMNH 34399 (1, dry); previous number SSB

3992 [883].

Paratypes: SBMNH 34400 (2, dry); previous number SSB

3993 [881,886].

California, Los Angeles Co., Santa Monica [=Los Angeles], Long Wharf Canyon [Potrero Canyon]; upper San Pedro Pleistocene [upper Pleistocene, Palos Verdes Sand or equiv- alent].

Remarks: Holotype (1 intermediate valve) illustrated in figs. 4-6; paratypes (1 head valve, 1 tail valve) illustrated in figs. 1-3, 7-9.

semiliratus, Dendrochiton

Berry, 1927: Proc. Malacol. Soc. London 17(4):160-163, text figs. 1-4.

Paratypes: SBMNH 34405 (2, dry); previous number SSB 5130 [1472].

Paratypes: SBMNH 34406 (2, dry); previous number SSB 5129 [1470].

Canada, British Columbia, Vancouver Id., Departure Bay.

Remarks: SBMNH 34405 (1 articulated, 1 disarticulated), illustrated in text figs. 1-4; SBMNH 34406 articulated. Holotype CAS 064257.

sonorana, Stenoplax {Maugerella) conspicua

Berry, 1956: Leafl. Malacol. l(13):73-74.

Holotype: SBMNH 34427 (1, dry); previous number SSB 15131.

Paratypes: SBMNH 34428 (14, dry); previous number SSB 14949 [2230].

Paratypes: SBMNH 34429 (4, dry); previous number SSB 17359 [2233].

Paratypes: SBMNH 34430 (4, dry); previous number SSB [2231].

Paratypes: SBMNH 34431 (11, dry); previous number SSB [2232].

Mexico, Sonora, Puerto Penasco, W end of long bight; under stones on reef.

Remarks: All specimens articulated except SBMNH 34429 and 34431 where 1 specimen in each lot is disarticulated.

stearnsii, Ischnochiton

Dale, 1902: Proc. U.S. Natl. Mus. 24(1264):557-558.

Syntype: SBMNH 34459 (-, dry); previous number SSB 16974.

California, San Francisco Co., off Farallon Ids.; 37°23'N, 123°08'W; 391 fm [715 m]; clay; USFC Sta. 3104.

Remarks: S. S. Berry label “holotype fragment”; 1 intermediate valve fragment, ex USNM 109024.

subtilis, Lepidozona

Berry, 1956: Leafl. Malacol. 1(13):74.

Holotype: SBMNH 34432 (1, dry); previous number SSB 15132.

Paratypes: SBMNH 34434 (33, dry); previous number SSB 14962 [2239].

Mexico, Sonora, Puerto Penasco, W end of long bight; under stones in pools on reef; mid-tide.

Paratypes: SBMNH 34433 (13, dry); previous number SSB 14962 [2238].

Mexico, Sonora, W shore of Bahia Punta Penasco; under stones near low-tide level.

Remarks: All specimens articulated.

thamnoporus, Mopalia {Dendrochiton)

Berry, 1911: Proc. Acad. Natur. Sci. Philadelphia 63(1911): 487-490.

Lectotype: SBMNH 34370 (1, dry); previous number SSB 4101 [8a].

Paralectotypes: SBMNH 3437 1 (9, dry); previous number SSB 5048.

California, Monterey Co., off Monterey; 15 fm [27 m].

Remarks: Lectotype disarticulated, missing valves 3 and 4. Paralectotypes all specimens articulated, several valves cracked. Lectotype designated: Berry, S. S. 1919. Proc. Cal- if. Acad. Sci. (4)9(1 ):4-5, figs. 1-2. Name emended from Mopalia thamnopora in lectotype designation.

willetti, Ischnochiton {Lepidozona)

Berry, 1917: Proc. Calif. Acad. Sci. (4)7(10):236-238, text figs. 1, 2.

Holotype: SBMNH 34372 (1, dry); previous number SSB 3700 [159].

Paratypes: SBMNH 34492 (7, dry); previous number SSB 3699 [210].

Alaska, Forrester Id.; 15-20 fm [27-37 m].

Remarks: All specimens articulated. Holotype illustrated in text figs. 1, 2.

Callistoplacidae

meridionalis, Callistochiton antiquus

Ashby, 1919: Trans. Roy. Soc. So. Australia 43:400. Paratypes: SBMNH 34456 (7, dry); previous number SSB 4551.

Australia, South Australia, St. Vincent’s Gulf, Marino. Remarks: All specimens articulated. Holotype reported in the South Australia Museum, Adelaide (not verified in this study).

Mopaliidae

aepynotus, Callistochiton

Dall, 1919: Proc. U.S. Natl. Mus. 55(2283):51 1.

Syntype: SBMNH 34466 (-, dry); previous number SSB 14560.

Washington, Puget Sound; 48°H'30"N, 122°48'W; 37 fm [68 m]; rocky bottom; USFC Sta. 3593.

Remarks: S. S. Berry label “holotype fragment.” ex USNM 225448.

chacei, Mopalia (Figure 1)

Berry, 1919: Lorquinia 2(6):46.

Lectotype (herein): SBMNH 34393 (1, 70% EtOH); previous number SSB 4129 [118].

Paralectotypes: SBMNH 35048 (6, dry); previous number SSB [767].

Type locality: California, San Diego Co., La Jolla; Coll. S. S. Berry, 18 Feb. 1913.

Paralectotype: SBMNH 35049 (1, dry); previous number SSB 3968 [745].

California, Los Angeles Co., Santa Monica, 3.7 mi [6 km] N of Long Wharf.

Remarks: S. S. Berry specimen label and collection catalog designated SBMNH 34393 as the “type”; lectotype specimen articulated on body and girdle; SBMNH 35048 3 speci- mens articulated, 3 disarticulated; SBMNH 35049 dis- articulated, several valves broken.

cirrata, Mopalia

Berry, 1919: Lorquinia 2(6):45-46.

Holotype (monotypy): SBMNH 34387 (1, dry); previous num- ber SSB [642].

Alaska, Icy Strait, Dundas Bay [bearing from Pt. Wimbleton,

Page 8

The Veliger, Vol. 33, Suppl. 1

S 20° W, 0.4 mi (0.6 km)]; 6.5-9 fm [12-16 mj; USFC Sta. 4263.

Remarks: 2 valves disarticulated, remainder articulated on gir- dle. S. S. Berry did not designate a holotype but his card file indicates only one specimen was in the type lot. The only other lot in the Berry collection was collected after the date of publication of the original species description.

cithara, Mopalia

Berry, 1951: Proc. Malacol. Soc. London 28(6):220, 222-224, pi. 26, figs. 16, 18, text figs. 1-8.

Holotype: .SBMNH 34422 (1, 70% EtOH); previous number SSB 13013 [2102].

Canad.\, British Columbia, Vancouver Id., just inside Clay- oquot Sound, SE shore of Round Id.; on boulders at —0.5 ft [—0.15 m] tide.

Remarks: 2 disarticulated valves, remainder articulated on body; head valve missing.

egretta, Mopalia

Berry, 1919: Lorquinia 2(6):45.

Lectotype: SBMNH 34384 (1, dry); previous number SSB

3939 [281].

Paralectotype: SBMNH 34385 (1, 70% EtOH); previous num- ber SSB [284].

Paralectotype: SBMNH 34386 (1, dry); previous number SSB

3940 [640].

Paralectotype: SBMNH 34448 (1, dry); previous number SSB 12645 [2073].

.Alaska, Forrester Id.; 20 fm [37 m].

Remarks: Lectotype designated: Berry, S. S. 1919. Proc. Calif. •\cad. Sci. (4)9(1): 13- 17; lectotype (articulated) illustrated in pi. 6, fig. 1; SBMNH 34385 illustrated in pi. 6, fig. 2; SBMNH 34386 illustrated in pi. 7, figs. 1-6. SBMNH 34448 is not listed in the lectotype designation but the S. S. Berry specimen label and corresponding collection catalog refer to it as a “paratype.”

lactica, IChaetopleura

Dale. 1919: Proc. U.S. Natl. Mus. 55(2283):509-510.

Syntype: SBMNH 34465 (1, dry); previous number SSB 14559.

California, Los Angeles Co., Santa Catalina Id., Catalina I larbor.

Remarks: Specimen articulated, ex USNM 172900.

lionolus, Mopalia imporcata

PiLSBRY, 1918: Nautilus 31(4):126.

Paratype: SBMNH 35084 (1, dry); previous number SSB [1812].

California, Los .\ngeles Co., White’s Point.

Remarks: ex H. N. Lowe No. 1542. .All but one valve artic- ulated on girdle. Holotype ANSP 1 17952.

pacifica, Placiphorella (Figure 2)

Berry, 1919: Lorquinia 2(6):46.

Lectotype (herein): SBMNH 34394 (1, dry & 70% EtOH); previous number SSB [92].

Type locality: Alaska, Prince of Wales Id., Kasaan Bay, true bearing from center of Round Id., S 10° W at 0.4 mi [0.6 km]; 95-98 fm [174-179 m]; USFC Sta. 4245; Coll. 1 1 July 1903, Steamer Albatross.

Remarks: S. S. Berry specimen label and collection catalog designated this as the “Type.” Specimen disarticulated, body in alcohol.

phorminx, Mopalia

Berry, 1919: Lorquinia 2(6):46.

Lectotype: SBMNH 34388 (1, dry); previous number SSB 1769 [549].

California, Monterey Co., Monterey Bay, ledge off Pt. Pinos, on a rock fragment; “depth probably 200 feet” [61 m].

Remarks: .All but 2 valves articulated. Lectotype designated: Berry, S. S. 1919. Proc. Calif. Acad. Sci. (4)9(1):10-13, pi. 3, figs. 1, 2, pi. 4, figs. 1-3, pi. 5, figs. 1-3.

riifa, Placiphorella

Berry, 1917: Proc. Calif. Acad. Sci. (4)7(10):241-244, text figs. 3, 4.

Holotype: SBMNH 34373 (1, 70% EtOH); previous number SSB 1411 [240].

Paratypes: SBMNH 34374 (3, dry); previous number SSB 3836 [242].

Paratype: SBMNH 34376 (1 , dry); previous number SSB 3837 [242a].

.Alaska, Forrester Id.; 15-20 fm [27-37 m].

Paratype: SBMNH 34375 (1 , dry); previous number SSB 3836 [245].

.Alaska, Forrester Id.; 25 fm [46 m].

Remarks: .All specimens articulated except SBMNH 34376. Holotype illustrated in text fig. 3; SBMNH 34376 illustrated in text fig. 4.

tasmanica, Kopionella

Ashby, 1920: Trans. Roy. Soc. So. Australia 44:268-271.

Syntype: SBMNH 34454 (1, dry); previous number SSB 6731.

Australia, Tasmania, D’ Entrecasteaux Channel, Wood- bridge.

Remarks: Specimen articulated, smashed.

Chitonidae

corypheus, Chiton

Hedley & Hull, 1912: Proc. Linnean Soc. New So. Wales 37(2):277-278.

Syntypes: SBMNH 34472 (2, dry); previous number SSB 5524.